Distinguished for his contributions to neurobiology. After graduating in Electrical Engineering he became involved in a study of synaptic transmission at autonomic nerve-smooth muscle junctions which culminated in an authoritative monograph written in 1969 at the invitation of the British Physiological Society. Subsequently, at the University of Sydney, he has been involved in analysing the factors which regulate the release of noradrenaline and acetylcholine from nerve terminals, and the mechanisms which determine the sites at which nerve terminals form synapses, both during development and after nerve lesions. His vigorous approach, technical expertise, and clarity in expressing and interpreting his observations have attracted a number of high quality graduate students, and in all aspects of his research he continues to contribute fundamental and very significant observations which undoubtedly assist the clarification of our understanding of synaptic mechanisms.